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and membrane trafficking\uff09\u3001Cantley\u535a\u58eb\uff08PI3 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<em><\/em><\/p>\n\n\n\n<p><strong>\u30fb<\/strong><strong>\u300c\u82e5\u624b\u80b2\u6210\u306e\u70ba\u306e\u5834\u300d\u3068\u3057\u3066\u306e\u65e5\u672c\u751f\u5316\u5b66\u4f1a\u5927\u4f1a<\/strong><\/p>\n\n\n\n<p>\u767a\u8868\u304c\u6bcd\u56fd\u8a9e\u3067\u304d\u3061\u3093\u3068\u6b63\u78ba\u306b\u767a\u8868\u3067\u304d\u308b\u3053\u3068\u304c\u57fa\u672c\u3067\u3042\u308a\u3001\u305d\u308c\u304c\u3067\u304d\u305f\u5f8c\u306b\u82f1\u8a9e\u3067\u767a\u8868\u3059\u308c\u3070\u826f\u3044\u3068\u79c1\u306f\u8003\u3048\u3066\u3044\u308b\u3002\u305d\u306e\u70ba\u306b\u65e5\u672c\u8a9e\u3067\u304d\u3061\u3093\u3068\u3057\u305f\u767a\u8868\u304c\u3067\u304d\u308b\u5834\u3068\u3057\u3066\u3001\u767a\u8868\u306f\u65e5\u672c\u8a9e\u306b\u3057\u305f\u3002\u4f46\u3057\u30b9\u30e9\u30a4\u30c9\u306f\u5168\u3066\u82f1\u8a9e\u3067\u8a18\u8f09\u3059\u308b\u3053\u3068\u306b\u3057\u3066\u3044\u305f\u70ba\u3001\u6d77\u5916\u304b\u3089\u306e\u62db\u5f85\u8005\u3082\u53c2\u52a0\u3057\u3066\u71b1\u5fc3\u306b\u30b9\u30e9\u30a4\u30c9\u3092\u898b\u3066\u8cea\u554f\u3059\u308b\u5834\u9762\u3082\u591a\u304f\u898b\u3089\u308c\u305f\u3002<\/p>\n\n\n\n<p>\u5b66\u751f\u3001\u5927\u5b66\u9662\u751f\u3092\u542b\u3081\u30662637\u540d\u306e\u53c2\u52a0\u3067\u3053\u308c\u3082\u5f53\u6642\u306e\u751f\u5316\u5b66\u4f1a\u3067\u306f\u53f2\u4e0a\u6700\u9ad8\u3060\u3063\u305f\u3002\u5b66\u751f\u304c\u591a\u304f\u306e\u5148\u751f\u65b9\u304b\u3089\u8cea\u554f\u3092\u53d7\u3051\u7dca\u5f35\u3057\u306a\u304c\u3089\u7b54\u3048\u3066\u3044\u308b\u5834\u9762\u306f\u5fae\u7b11\u307e\u3057\u304b\u3063\u305f\u3002\u4eca\u56de\u53c2\u52a0\u3057\u305f\u5b66\u751f\u306f\u5927\u5909\u9762\u767d\u304b\u3063\u305f\uff64\u6765\u5e74\u306f\u82e5\u3044\u5b66\u751f\u3092\u9023\u308c\u3066\u6765\u305f\u3044\u3068\u3044\u3046\u58f0\u304c\u805e\u3053\u3048\u305f\u3002\u5b66\u751f\u306e\u53c2\u52a0\u306b\u4f34\u3063\u3066\u3001\u5404\u30dd\u30b9\u30bf\u30fc\u306b\u306f\u591a\u304f\u306e\u4eba\u304c\u96c6\u307e\u308a\u71b1\u5fc3\u306b\u8a0e\u8b70\u3092\u3057\u3066\u304a\u308a\u3001\u71b1\u6c17\u306b\u3042\u3075\u308c\u3066\u3044\u305f\u3002<\/p>\n\n\n\n<p><strong>\u30fb<\/strong><strong>\u53e3\u982d\u767a\u8868\u3092\u5927\u5e45\u306b\u5897\u52a0<\/strong><\/p>\n\n\n\n<p>\u30d7\u30ed\u30b0\u30e9\u30e0\u7de8\u96c6\u59d4\u54e1\u4f1a\u3067\u53e3\u982d\u767a\u8868\uff08\u65e5\u672c\u8a9e\uff09\u3092\u5927\u5e45\u306b\u5897\u3084\u3057\u305f\u3002\u53e3\u982d\u767a\u8868\u3092\u30c6\u30fc\u30de\u5225\u306b\u307e\u3068\u3081\u3066\u30ef\u30fc\u30af\u30b7\u30e7\u30c3\u30d7\u3068\u3057\u3066\u8a2d\u3051\u305f\u3002\u3067\u304d\u308b\u3060\u3051\u591a\u304f\u306e\u82e5\u3044\u7814\u7a76\u8005\u3084\u5927\u5b66\u9662\u751f\u306e\u4f1a\u54e1\u306b\u53e3\u982d\u3067\u767a\u8868\u3057\u3066\u9802\u304f\u6a5f\u4f1a\u3092\u5927\u5e45\u306b\u5897\u3084\u3057\u305f\u3002\u3053\u306e10\u5e74\u7a0b\u30dd\u30b9\u30bf\u30fc\u306b\u3070\u304b\u308a\u6163\u308c\u3066\u767a\u8868\u304c\u304d\u3061\u3093\u3068\u3067\u304d\u306a\u3044\u5b66\u751f\u304c\u591a\u304f\u306a\u3063\u305f\u70ba\u3001\u30c8\u30ec\u30fc\u30cb\u30f3\u30b0\u3082\u517c\u306d\u305f\u3002\u81f3\u308b\u6240\u3067\u7dca\u5f35\u3057\u306a\u304c\u3089\u3082\u767a\u8868\u306e\u70ba\u306e\u7df4\u7fd2\u3092\u3057\u3066\u3044\u308b\u6b21\u4e16\u4ee3\u3092\u62c5\u3046\u82e5\u3044\u4eba\u305f\u3061\u306e\u59ff\u306f\u5fae\u7b11\u307e\u3057\u3044\u3082\u306e\u3067\u3042\u3063\u305f\u3002<\/p>\n\n\n\n<p><strong>\u30fb<\/strong><strong>\u767b\u9332\u8cbb\u3001\u53c2\u52a0\u8cbb\u3001\u61c7\u89aa\u4f1a\u8cbb\u3092\u5927\u5e45\u306b\u4f4e\u304f\u3057\u305f<\/strong><\/p>\n\n\n\n<p>\u958b\u50ac\u8005\u5074\u3068\u3057\u3066\u8cc7\u91d1\u3092\u96c6\u3081\u308b\u52aa\u529b\u3092\u3057\u305f\u3002\u4e8b\u524d\u767b\u9332\u8cbb\u3092\u9662\u751f\u306f2\u5343\u5186\u3001\u5b66\u751f\u306f\u7121\u6599\u3068\u3057\u3001\u61c7\u89aa\u4f1a\u8cbb\u3082\u601d\u3044\u5207\u3063\u3066\u5b9f\u8cea1\u4e07\u5186\u30922\u5343\u5186\u3068\u3057\u305f\u3053\u3068\u3082\u82e5\u624b\u3092\u60f9\u304d\u3064\u3051\u305f\u4e00\u56e0\u3067\u3042\u308d\u3046\u3002\u7f8e\u5473\u3057\u3044\u6599\u7406\u3092\u5b89\u304f\u63d0\u4f9b\u3057\uff64\u82e5\u624b\u304b\u3089\u5e74\u914d\u307e\u3067\u4f1a\u54e1\u306e\u4eba\u7684\u4ea4\u6d41\u304c\u3088\u308a\u6d3b\u767a\u3068\u306a\u308b\u3088\u3046\u306b\u5fc3\u304c\u3051\u305f\u3002\u5927\u4f1a\u306e\u958b\u50ac\u306b\u3042\u305f\u308a\u65e5\u672c\u88fd\u85ac\u56e3\u4f53\u9023\u5408\u4f1a\u3092\u306f\u3058\u3081\u3068\u3057\u3066\u52a9\u6210\u8ca1\u56e3\u3088\u308a\u591a\u5927\u306a\u3054\u63f4\u52a9\u3092\u9802\u3044\u305f\u3002<\/p>\n\n\n\n<p><strong>\u30fb<\/strong><strong>\u4f01\u696d\u304b\u3089\u306e\u30b5\u30dd\u30fc\u30c8\u3092\u7a4d\u6975\u7684\u306b\u63a8\u9032<\/strong><\/p>\n\n\n\n<p>\u4f01\u696d\u304b\u3089\u306e\u30b5\u30dd\u30fc\u30c8\u306b\u3088\u308b\u30d6\u30fc\u30b9\u306e\u6570\u306f450\u4ee5\u4e0a\u3068\uff64\u7686\u3067\u58f0\u304c\u3051\u3092\u884c\u3063\u305f\u7d50\u679c\u591a\u304f\u306a\u3063\u305f\u30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Kornberg\u5148\u751f\u306b\u3001\u307e\u305f\u65e9\u77f3\u4fee\u5148\u751f\u306b\u3082\u30de\u30b9\u30bf\u30fc\u30ba\u30ec\u30af\u30c1\u30e3\u30fc\u3092\u304a\u9858\u3044\u3057\u305f\u3002\u4e00\u822c\u6f14\u984c2303\u6f14\u984c\uff08\u30ef\u30fc\u30af\u30b7\u30e7\u30c3\u30d7901\u6f14\u984c\uff09\u53ca\u307324\u30c6\u30fc\u30de\u306b\u3064\u3044\u3066\u306e\u30b7\u30f3\u30dd\u30b8\u30a6\u30e0\u3092\u958b\u50ac\u3057\u305f\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\"><strong>\u65e5\u672c\u751f\u5316\u5b66\u4f1a\u306e\u751f\u547d\u79d1\u5b66\u3078\u306e\u3053\u308c\u307e\u3067\u306e\u8ca2\u732e<\/strong><\/h3>\n\n\n\n<p>\u65e5\u672c\u751f\u5316\u5b66\u4f1a\u306f\u65e5\u672c\u306e\u4e2d\u5fc3\u7684\u306a\u5b66\u4f1a\u306e\u4e00\u3064\u3068\u3057\u3066\u751f\u547d\u79d1\u5b66\u306e\u767a\u5c55\u306b\u8ca2\u732e\u3057\u3066\u304d\u305f\u3053\u3068\u306f\u660e\u3089\u304b\u3067\u3042\u308b\u3068\u79c1\u306f\u8003\u3048\u308b\u3002\u305d\u306e\u65b9\u6cd5\u8ad6\u306f\u5f53\u521d\u306f\u30c7\u30ab\u30eb\u30c8\u306e\u8981\u7d20\u9084\u5143\u8ad6\u7684\u306a\u65b9\u5411\u304c\u4e2d\u5fc3\u3067\u306f\u3042\u3063\u305f\u304c\u4fef\u77b0\u7684\u306a\u7d71\u5408\u8ad6\u306a\u89e3\u6790\u3092\u3082\u542b\u3081\u3066\u9032\u3081\u3066\u304d\u305f\u91cd\u539a\u306a\u7814\u7a76\u8005\u5c64\u3092\u6301\u3064\u65e5\u672c\u751f\u5316\u5b66\u4f1a\u306e\u8ca2\u732e\u306f\u8a55\u4fa1\u3059\u3079\u304d\u3068\u8003\u3048\u308b\u3002<\/p>\n\n\n\n<p>\u3055\u3066\u3001\u56fd\u969b\u7684\u306b\u5927\u304d\u304f\u8a8d\u77e5\u3055\u308c\u3066\u3044\u308b\u65e5\u672c\u751f\u5316\u5b66\u4f1a\u306e\u679c\u305f\u3059\u5f79\u5272\u306f\u4eca\u307e\u3067\u306e\u3082\u306e\u306e\u307f\u3067\u5341\u5206\u3067\u3042\u3063\u305f\u3060\u308d\u3046\u304b\uff1f\u3000\u79c1\u306f\u73fe\u5728\u306e\u5b66\u4f1a\u306e\u69cb\u6210\u54e1\u3068\u5b9f\u529b\u3092\u8003\u3048\u3066\u5341\u5206\u306b\u5927\u304d\u304f\u5c55\u958b\u3059\u308b\u529b\u304c\u3042\u308b\u3068\u8003\u3048\u3066\u3044\u308b\u3002\u6b21\u306e\u9805\u3067\u73fe\u5728\u79c1\u9054\u304c\u5bfe\u5fdc\u3057\u306a\u3051\u308c\u3070\u306a\u3089\u306a\u3044\u8ab2\u984c\u306b\u3064\u3044\u3066\u8ff0\u3079\u3066\u3044\u304f\u3002<\/p>\n\n\n\n<h3 class=\"wp-block-heading\"><strong>\u4eba\u985e\u3001\u5730\u7403\u306e\u5371\u6a5f\u2014\u2014\u30e9\u30a4\u30d5\u30b5\u30a4\u30a8\u30f3\u30b9\u306e\u9032\u5c55\u306b\u3088\u308b\u5730\u7403\u74b0\u5883\u306e\u7834\u58ca<\/strong><strong><\/strong><\/h3>\n\n\n\n<h4 class=\"wp-block-heading\"><strong>\u2014\u2014\u65e5\u672c\u751f\u5316\u5b66\u4f1a\u306e\u7f6e\u304b\u308c\u3066\u3044\u308b\u91cd\u8981\u306a\u7acb\u5834\u3092\u7406\u89e3\u3057\u3088\u3046\u2014\u2014<\/strong><strong><\/strong><\/h4>\n\n\n\n<p>\u3055\u3066\u3082\u3046\u4e00\u5ea6\u73fe\u5728\u79c1\u9054\u304c\u7f6e\u304b\u308c\u3066\u3044\u308b\u74b0\u5883\u3092\u8003\u3048\u3066\u307f\u308b\u3002\u73fe\u5728\u9032\u3081\u3066\u3044\u308b\u30e9\u30a4\u30d5\u30b5\u30a4\u30a8\u30f3\u30b9\u306e\u7814\u7a76\u306e\u9032\u5c55\u306f\u6211\u3005\u306e\u4e88\u60f3\u3092\u9065\u304b\u306b\u8d85\u3048\u3066\u304a\u308a\uff64\u3053\u306e\u307e\u307e\u7d9a\u3051\u3066\u3044\u304f\u3068\u5730\u7403\u74b0\u5883\u3082\u542b\u3081\u3066\u4eba\u9593\u306e\u5b58\u5728\u305d\u306e\u3082\u306e\u304c\u8105\u304b\u3055\u308c\u308b\uff08\u88681\u53c2\u7167\uff09\u3002<\/p>\n\n\n\n<div class=\"wp-block-group stripe-table is-layout-constrained wp-block-group-is-layout-constrained\">\n<p><strong>\u88681\u3000\u30e9\u30a4\u30d5\u30b5\u30a4\u30a8\u30f3\u30b9\u7814\u7a76\u306e\u4e88\u60f3\u3092\u8d85\u3048\u305f\u767a\u5c55\u306b\u3088\u308b\u4eba\u985e\u3001\u5730\u7403\u306e\u5371\u6a5f\u2014\u2014\u751f\u547d\u79d1\u5b66\u306e\u767a\u5c55\u306e\u529f\u7f6a\u2014\u2014<\/strong><sup>5)<\/sup><\/p>\n\n\n\n<figure class=\"wp-block-table is-style-vk-table-border-stripes\"><table class=\"has-fixed-layout\"><tbody><tr><td><strong>\u79d1\u5b66\u6280\u8853\u306e\u8105\u5a01\u3000\u5149\u3068\u5f71\u3068\u95c7<\/strong><\/td><\/tr><tr><td>&nbsp;<\/td><\/tr><tr><td>\u30a8\u30cd\u30eb\u30ae\u30fc\uff1a\u7269\u8cea\u306f\u30a8\u30cd\u30eb\u30ae\u30fc\u3001\u539f\u5b50\u30a8\u30cd\u30eb\u30ae\u30fc\u306e\u5e73\u548c\u5229\u7528\u3068\u539f\u5b50\u7206\u5f3e<\/td><\/tr><tr><td>\u30c0\u30a4\u30ca\u30de\u30a4\u30c8\uff1a\u5efa\u8a2d\u3001\u571f\u6728\u5de5\u4e8b\u3001\u7206\u5f3e\u3001\u6226\u4e89\u306e\u6b66\u5668<\/td><\/tr><tr><td>\u5b87\u5b99\u3001\u5de8\u5927\u5b87\u5b99\u30d7\u30ed\u30b8\u30a7\u30af\u30c8\uff1a\u5b87\u5b99\u958b\u767a\u3001\u5b87\u5b99\u30b9\u30c6\u30fc\u30b7\u30e7\u30f3\u3001\u5b87\u5b99\u57fa\u5730<\/td><\/tr><tr><td>\u7dd1\u5316\uff1a\u690d\u7269\u3068\u52d5\u7269\u3001CO<sub>2<\/sub>-O<sub>2<\/sub>\u95a2\u9023\u3001\u8d85\u9ad8\u97f3\u901f\u30b8\u30a7\u30c3\u30c8\u6a5f\uff08\u30b3\u30f3\u30b3\u30eb\u30c9\uff09\u306b\u3088\u308b\u30a2\u30de\u30be\u30f3\u306e\u68ee\u6797\u7834\u58ca<\/td><\/tr><tr><td>\u6d77\u6d0b\uff1a\u6d77\u6d0b\u6c5a\u67d3\u3001\u9b5a\u306e\u6c5a\u67d3\u3001\u5fae\u7c92\u5b50\u30d7\u30e9\u30b9\u30c1\u30c3\u30af<\/td><\/tr><tr><td>CO<sub>2<\/sub>\uff1a\u6d77\u6d0b\u4e2d\u306eCO<sub>2<\/sub>\u84c4\u7a4d\u91cf<\/td><\/tr><tr><td>\u30d7\u30e9\u30b9\u30c1\u30c3\u30af\uff1a\u30da\u30c3\u30c8\u30dc\u30c8\u30eb\u3001\u5730\u7403\u306e\u6c5a\u67d3<\/td><\/tr><tr><td>\u30b2\u30ce\u30e0\u30d7\u30ed\u30b8\u30a7\u30af\u30c8\u3001\u907a\u4f1d\u5b50\u3001\u30a6\u30a4\u30eb\u30b9\uff1a\u907a\u4f1d\u5b50\u6539\u5909\u3001\u611f\u67d3\u6027\u306e\u5236\u5fa1\u2026\u6bd2\u6027\u306e\u9664\u53bb\u3001\u5f37\u6bd2\u682a\u306e\u7523\u751f\u306a\u3069\u81ea\u7531\u306a\u5909\u63db<\/td><\/tr><tr><td>\u6c17\u5019\u5909\u52d5\u3001\u5730\u7403\u6e29\u6696\u5316\u3001\u98df\u7ce7\u5371\u6a5f\u3001\u56fd\u5bb6\u9593\u306e\u6226\u4e89\u52c3\u767a<\/td><\/tr><\/tbody><\/table><\/figure>\n<\/div>\n\n\n\n<div style=\"height:49px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<p>\u5730\u7403\u4e0a\u306b\u751f\u80b2\u3059\u308b\u751f\u7269\u306f\u4eba\u985e\u3060\u3051\u3067\u306f\u306a\u3044\u3002\u9032\u5316\u306e\u904e\u7a0b\u3067\u591a\u304f\u306e\u751f\u7269\u304c\u751f\u307e\u308c\u3001\u305d\u306e\u4e00\u3064\u3068\u3057\u3066\u4eba\u9593\u304c\u51fa\u3066\u304d\u305f\u3002\u30e9\u30a4\u30d5\u30b5\u30a4\u30a8\u30f3\u30b9\u306e\u6025\u6fc0\u306a\u9032\u6b69\u3092\u8e0f\u307e\u3048\uff64\u4eba\u9593\u3068\u3044\u3046\u751f\u7269\u304c\uff64\u4ed6\u306e\u751f\u7269\u3068\u5171\u5b58\u3057\u306a\u304c\u3089\uff64\u5730\u7403\uff64\u4e16\u754c\uff64\u5730\u57df\u3067\uff64\u81ea\u7136\u74b0\u5883\u306b\u56f2\u307e\u308c\uff64\u8c4a\u304b\u306b\u751f\u5b58\u3059\u308b\u3053\u3068\u304c\u5fc5\u9808\u306a\u72b6\u6cc1\u306b\u306a\u3063\u3066\u304d\u3066\u3044\u308b\u3002<strong><\/strong><\/p>\n\n\n\n<p>\u4eba\u9593\u3068\u3044\u3046\u751f\u7269\u304c\u751f\u307e\u308c\u80b2\u3063\u305f\u300c\u5730\u7403\u306e\u5b58\u7d9a\u300d\u3092\u8003\u3048\u306a\u304c\u3089\uff64\u4ed6\u306e\u751f\u7269\u3068\u3044\u304b\u306b\u5171\u5b58\u3057\u3066\u3044\u304f\u304b\u306b\u3064\u3044\u3066\u4eba\u985e\u306e\u300c\u751f\u5b58\u300d\u3092\u6b63\u9762\u304b\u3089\u63a1\u308a\u4e0a\u3052\u3056\u308b\u3092\u5f97\u305a\uff64\u65e2\u5b58\u306e\u79d1\u5b66\u65b9\u6cd5\u8ad6\u306b\u62d8\u6ce5\u3059\u308b\u9650\u308a\uff64\u3053\u306e\u72b6\u6cc1\u306b\u5bfe\u5fdc\u3067\u304d\u306a\u3044\u3053\u3068\u306f\u660e\u767d\u3067\u3042\u308b\u3002\u305d\u306e\u305f\u3081\u306b\uff64\u5168\u3066\u306e\u79d1\u5b66\u306f\uff64\u300c\u751f\u5b58\u300d\u554f\u984c\u3092\u524d\u306b\u3057\u3066\u53e4\u5178\u7684\u306a\u9589\u9396\u7684\u67a0\u7d44\u307f\u3092\u5909\u3048\u308b\u5fc5\u8981\u304c\u3042\u308b\u3060\u308d\u3046\u3002\u81ea\u7136\u79d1\u5b66\u3068\u793e\u4f1a\u79d1\u5b66\u3068\u3044\u3046\u533a\u5225\u3059\u3089\u3082\u306a\u304f\u3057\u3066\u2500\u2500\u533b\u5b66\uff64\u7269\u7406\u5b66\uff64\u5316\u5b66\uff64\u5de5\u5b66\u306a\u3069\u306f\u3082\u3061\u308d\u3093\u306e\u3053\u3068\uff64\u7d4c\u6e08\u5b66\uff64\u6cd5\u5f8b\u5b66\uff64\u653f\u6cbb\u5b66\u306a\u3069\u3082\u542b\u3081\u3066\u2500\u2500\u3042\u3089\u3086\u308b\u9818\u57df\u304b\u3089\u7dcf\u5408\u7684\u306b\u300c\u751f\u5b58\u300d\u554f\u984c\u306b\u53d6\u308a\u7d44\u307e\u3056\u308b\u3092\u5f97\u306a\u3044\u72b6\u6cc1\u3067\u3042\u308b\uff08\u6b66\u898b\u592a\u90ce\u535a\u58eb\u306b\u3088\u308b\u300c\u751f\u5b58\u79d1\u5b66\u300d\u306e\u63d0\u5531<sup>1-3\u3001 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